About the Authors

Yang Gao

Affiliations Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, University of Chinese Academy of Sciences, Beijing, China

Huan Zhang

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Qiang Gao

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Lingling Wang

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Fuchong Zhang

fuchongz@yahoo.com.cn (FZ); lshsong@qdio.ac.cn (LS)

Affiliation Ocean and Fishery Sciences Research Institute of Hebei Province, Qinhuangdao, China

Vinu S. Siva

Affiliations Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, University of Chinese Academy of Sciences, Beijing, China

Zhi Zhou

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Linsheng Song

fuchongz@yahoo.com.cn (FZ); lshsong@qdio.ac.cn (LS)

Affiliation Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Shicui Zhang

Affiliation Ocean University of China, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LS FZ YG. Performed the experiments: YG HZ VSS ZZ. Analyzed the data: YG QG. Contributed reagents/materials/analysis tools: YG LW HZ SZ. Wrote the paper: YG HZ LW FZ LS.