About the Authors

Myles Benjamin Walton

Affiliation Department of Musculoskeletal Biology and School of Veterinary Science, University of Liverpool, Neston, Wirral, United Kingdom

Emily Cowderoy

Affiliation Department of Musculoskeletal Biology and School of Veterinary Science, University of Liverpool, Neston, Wirral, United Kingdom

Duncan Lascelles

Affiliation Comparative Pain Research Laboratory and Center for Comparative Medicine and Translational Research, and College of Veterinary Medicine, North Carolina State University, Raleigh, North Carolina, United States of America

John F. Innes

j.f.innes@liv.ac.uk

Affiliation Department of Musculoskeletal Biology and School of Veterinary Science, University of Liverpool, Neston, Wirral, United Kingdom

Competing Interests

This work was performed during a period when MBW’s and EC’s salaries were funded by Pfizer Animal Health. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JFI DL MBW. Performed the experiments: MBW EC. Analyzed the data: MBW. Contributed reagents/materials/analysis tools: JFI. Wrote the paper: MBW EC DL JFI.