About the Authors

So-Hyeon Baek

Affiliation National Institute of Crop Science, Rural Development Administration, Iksan, Chonbuk, Korea

Woon-Chul Shin

Affiliation National Institute of Crop Science, Rural Development Administration, Iksan, Chonbuk, Korea

Hak-Seung Ryu

Affiliations Graduate School of Biotechnology, Kyung Hee University, Yongin, Gyeonggi, Korea, Crop Biotech Institute, Kyung Hee University, Yongin, Gyeonggi, Korea

Dae-Woo Lee

Affiliations Graduate School of Biotechnology, Kyung Hee University, Yongin, Gyeonggi, Korea, Crop Biotech Institute, Kyung Hee University, Yongin, Gyeonggi, Korea

Eunjung Moon

Affiliation Graduate School of East-West Medical Science, Kyung Hee University, Yongin, Gyeonggi, Korea

Chun-Sun Seo

Affiliation National Institute of Crop Science, Rural Development Administration, Iksan, Chonbuk, Korea

Eunson Hwang

Affiliation Graduate School of East-West Medical Science, Kyung Hee University, Yongin, Gyeonggi, Korea

Hyun-Seo Lee

Affiliation Laboratory of Genetics and Department of Microbiology, Chonbuk National University Medical School, Jeonju, Chonbuk, Korea

Mi-Hyun Ahn

Affiliation Laboratory of Genetics and Department of Microbiology, Chonbuk National University Medical School, Jeonju, Chonbuk, Korea

Youngju Jeon

Affiliation Laboratory of Genetics and Department of Microbiology, Chonbuk National University Medical School, Jeonju, Chonbuk, Korea

Hyeon-Jung Kang

Affiliation National Institute of Crop Science, Rural Development Administration, Iksan, Chonbuk, Korea

Sang-Won Lee

Affiliations Crop Biotech Institute, Kyung Hee University, Yongin, Gyeonggi, Korea, Department of Plant Molecular Systems Biotechnology, Kyung Hee University, Yongin, Gyeonggi, Korea

Sun Yeou Kim

Affiliation College of pharmacy, Gachon University, Incheon, Korea

Roshan D’Souza

Affiliation BDRD Research Institute, JINIS Biopharmaceuticals Inc., Wanju, Chonbuk, Korea

Hyeon-Jin Kim

Affiliation BDRD Research Institute, JINIS Biopharmaceuticals Inc., Wanju, Chonbuk, Korea

Seong-Tshool Hong

seonghong@chonbuk.ac.kr (STH); jjeon@khu.ac.kr (JSJ)

Affiliation Laboratory of Genetics and Department of Microbiology, Chonbuk National University Medical School, Jeonju, Chonbuk, Korea

Jong-Seong Jeon

seonghong@chonbuk.ac.kr (STH); jjeon@khu.ac.kr (JSJ)

Affiliations Graduate School of Biotechnology, Kyung Hee University, Yongin, Gyeonggi, Korea, Crop Biotech Institute, Kyung Hee University, Yongin, Gyeonggi, Korea

Competing Interests

RD and H. Kim are employees of BDRD Research Institute, JINIS Biopharmaceuticals, Inc. BDRD Research Institute, JINIS Biopharmaceuticals, Inc. helped to carry out in vivo efficacy assay of transgenic grains using a mouse model. All other authors declare no competing interest. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SB SYK SH JJ. Performed the experiments: SB WS HR DL EM CS EH HL MA YJ H. Kang SL RD H. Kim. Analyzed the data: SB HR SL SYK SH JJ. Wrote the paper: SB HR SL SYK SH JJ.