About the Authors

Ferry Cornelissen

Affiliations Department of Rheumatology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands, Department of Immunology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands

Patrick S. Asmawidjaja

Affiliations Department of Rheumatology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands, Department of Immunology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands

Adriana M. C. Mus

Affiliations Department of Rheumatology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands, Department of Immunology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands

Odilia Corneth

Affiliations Department of Rheumatology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands, Department of Immunology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands

Kristine Kikly

Affiliation Eli Lilly and Company, Indianapolis, Indiana, United States of America

Erik Lubberts

E.Lubberts@erasmusmc.nl

Affiliations Department of Rheumatology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands, Department of Immunology, Erasmus Medical Center Rotterdam, Rotterdam, The Netherlands

Competing Interests

Dr. Kikly is an employee of Eli Lilly and Company. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FC PA KK EL. Performed the experiments: FC PA AM OC EL. Analyzed the data: FC PA AM KK EL. Contributed reagents/materials/analysis tools: FC PA AM OC KK EL. Wrote the paper: FC KK EL.