About the Authors

Koichi Oishi

Affiliation Department of Advanced Medicine in Biotechnology and Robotics, Nagoya University Graduate School of Medicine, Nagoya, Aichi, Japan

Yoshitaka Miyamoto

Affiliation Department of Advanced Medicine in Biotechnology and Robotics, Nagoya University Graduate School of Medicine, Nagoya, Aichi, Japan

Hiroaki Saito

Affiliation Nagoya Research Laboratory, MEITO Sangyo Co., Ltd., Nagoya, Aichi, Japan

Katsutoshi Murase

Affiliation Nagoya Research Laboratory, MEITO Sangyo Co., Ltd., Nagoya, Aichi, Japan

Kenji Ono

Affiliation Department of Brain Functions, Division of Stress Adaptation and Protection, Research Institute of Environmental Medicine, Nagoya University, Nagoya, Aichi, Japan

Makoto Sawada

Affiliation Department of Brain Functions, Division of Stress Adaptation and Protection, Research Institute of Environmental Medicine, Nagoya University, Nagoya, Aichi, Japan

Masami Watanabe

Affiliation Center for Gene and Cell Therapy, Okayama University Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama, Okayama, Japan

Yasufumi Noguchi

Affiliation Department of Socio-environmental Design, Hiroshima International University, Kure, Hiroshima, Japan

Toshiyoshi Fujiwara

Affiliation Department of Gastroenterological Surgery, Okayama University Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama, Okayama, Japan

Shuji Hayashi

Affiliation Department of Advanced Medicine in Biotechnology and Robotics, Nagoya University Graduate School of Medicine, Nagoya, Aichi, Japan

Hirofumi Noguchi

noguch-h@cc.okayama-u.ac.jp

Affiliations Department of Advanced Medicine in Biotechnology and Robotics, Nagoya University Graduate School of Medicine, Nagoya, Aichi, Japan, Department of Gastroenterological Surgery, Okayama University Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama, Okayama, Japan

Competing Interests

The authors have read the journal's policy and have the following conflicts: Murase Katsutoshi has stock options and paid employment at Meito Sangyo, Inc. Hiroaki Saito has ownership of stocks and paid employment at Meito Sangyo, Inc. The following authors have no competing interests: Koichi Oishi, Yoshitaka Miyamoto, Kenji Ono, Makoto Sawada, Masami Watanabe, Yasufumi Noguchi, Toshiyoshi Fujiwara, Shuji Hayashi, Hirofumi Noguchi. MEITO Sangyo Co., Ltd. provided the nanoparticles used in this study (Trimethylamino dextran-coated, magnetic iron oxide nanoparticles (TMADM-03)). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HN. Performed the experiments: K. Oishi YM HS KM HN. Analyzed the data: YN HN. Contributed reagents/materials/analysis tools: K. Ono MS MW TF SH. Wrote the paper: K. Oishi HN.