About the Authors

Xiaofei Wang

Contributed equally to this work with: Xiaofei Wang, Shizhong Zhang

Affiliations National Key laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China

Shizhong Zhang

Contributed equally to this work with: Xiaofei Wang, Shizhong Zhang

Affiliations National Key laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China

Ling Su

Affiliations National Key laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China

Xin Liu

Affiliations National Key laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China

Yujin Hao

haoyujin@sdau.edu.cn

Affiliations National Key laboratory of Crop Biology, Shandong Agricultural University, Tai-An, Shandong, China, National Research Center for Apple Engineering and Technology, Shandong Agricultural University, Tai-An, Shandong, China, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XW SZ YH. Performed the experiments: XW LS. Analyzed the data: LS XL. Contributed reagents/materials/analysis tools: XL LS. Wrote the paper: XW SZ.