About the Authors

Melania Figueroa

Affiliation Forage Seed and Cereal Research Unit, Agricultural Research Service, U.S. Department of Agriculture, Corvallis, Oregon, United States of America

Stephen Alderman

Affiliation Forage Seed and Cereal Research Unit, Agricultural Research Service, U.S. Department of Agriculture, Corvallis, Oregon, United States of America

David F. Garvin

Affiliation Plant Science Research Unit and Department of Agronomy and Plant Genetics, Agricultural Research Service, U.S. Department of Agriculture, University of Minnesota. St. Paul, Minnesota, United States of America

William F. Pfender

pfenderw@onid.orst.edu

Affiliations Forage Seed and Cereal Research Unit, Agricultural Research Service, U.S. Department of Agriculture, Corvallis, Oregon, United States of America, Department of Botany and Plant Pathology, Oregon State University, Corvallis, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MF WFP. Performed the experiments: MF SA. Analyzed the data: MF WFP DFG SA. Contributed reagents/materials/analysis tools: WFP DFG SA. Wrote the paper: MF WFP.