About the Authors

Hiroaki Asai

Affiliation Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan

Hiroshi Fujiwara

yunarief@m.ehime-u.ac.jp (HF); yasukawa@m.ehime-u.ac.jp (MY)

Affiliations Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan, Department of Cell Growth and Tumor Regulation, Ehime University Proteo-Medicine Research Center, Ehime, Japan

Jun An

Affiliation Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan

Toshiki Ochi

Affiliations Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan, Princess Margaret Hospital, Ontario Cancer Institute, Ontario, Canada

Yukihiro Miyazaki

Affiliation Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan

Kozo Nagai

Affiliation Department of Pediatrics, Ehime University Graduate School of Medicine, Ehime, Japan

Sachiko Okamoto

Affiliation Center for Cell and Gene Therapy, Takara Bio Inc., Shiga, Japan

Junichi Mineno

Affiliation Center for Cell and Gene Therapy, Takara Bio Inc., Shiga, Japan

Kiyotaka Kuzushima

Affiliation Division of Immunology, Aichi Cancer Center, Aichi, Japan

Hiroshi Shiku

Affiliation Department of Cancer Vaccine and Immuno-Gene Therapy, Mie University Graduate School of Medicine, Mie, Japan

Hirofumi Inoue

Affiliation Department of Biochemistry and Molecular Genetics, Ehime University Graduate School of Medicine, Ehime, Japan

Masaki Yasukawa

yunarief@m.ehime-u.ac.jp (HF); yasukawa@m.ehime-u.ac.jp (MY)

Affiliations Department of Bioregulatory Medicine, Ehime University Graduate School of Medicine, Ehime, Japan, Department of Cell Growth and Tumor Regulation, Ehime University Proteo-Medicine Research Center, Ehime, Japan

Competing Interests

The authors except SO, JM and HS have declared that no competing interest exist. SO and JM are employee of Takara Bio, Inc.. HS is provided with research funding from Takara Bio, Inc.. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HF MY. Performed the experiments: HA HF JA TO YM KN. Analyzed the data: HA HF HS MY. Contributed reagents/materials/analysis tools: SO JM KK HI. Wrote the paper: HA HF MY.