About the Authors

Atle van Beelen Granlund

Contributed equally to this work with: Atle van Beelen Granlund, Arnar Flatberg

Affiliations Centre of Molecular Inflammation Research, Norwegian University of Science and Technology, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Arnar Flatberg

Contributed equally to this work with: Atle van Beelen Granlund, Arnar Flatberg

Affiliation Centre of Molecular Inflammation Research, Norwegian University of Science and Technology, Trondheim, Norway

Ann E. Østvik

Affiliations Centre of Molecular Inflammation Research, Norwegian University of Science and Technology, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastroenterology and Hepatology, St. Olav’s University Hospital, Trondheim, Norway

Ignat Drozdov

Affiliation Bering Limited, Richmond, United Kingdom

Bjørn I. Gustafsson

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastroenterology and Hepatology, St. Olav’s University Hospital, Trondheim, Norway

Mark Kidd

Affiliation Department of Surgery, Section of Gastroenterology, Yale School of Medicine, New Haven, Connecticut, United States of America

Vidar Beisvag

Affiliation Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Sverre H. Torp

Affiliations Department of Laboratory Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Pathology, St. Olav’s University Hospital, Trondheim, Norway

Helge L. Waldum

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastroenterology and Hepatology, St. Olav’s University Hospital, Trondheim, Norway

Tom Christian Martinsen

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastroenterology and Hepatology, St. Olav’s University Hospital, Trondheim, Norway

Jan Kristian Damås

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Infectious Diseases, St. Olav’s University Hospital, Trondheim, Norway

Terje Espevik

Affiliations Centre of Molecular Inflammation Research, Norwegian University of Science and Technology, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Arne K. Sandvik

arne.sandvik@ntnu.no

Affiliations Centre of Molecular Inflammation Research, Norwegian University of Science and Technology, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastroenterology and Hepatology, St. Olav’s University Hospital, Trondheim, Norway

Competing Interests

Ignat Drozdov is employed by Bering Limited. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Contributed substantially in results interpretation: TE JKD MK BG AKS. Conceived and designed the experiments: HW AKS ABG AF BG MK JKD TE. Performed the experiments: VB ABG ST. Analyzed the data: AF ABG ID MK TE JKD AKS. Contributed reagents/materials/analysis tools: AEØ TCM AKS HW BG TE. Wrote the paper: ABG AKS AF ID.