About the Authors

Keiji Ozeki

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Satoshi Tanida

stanida@med.nagoya-cu.ac.jp (ST); shigeki@m.ehime-u.ac.jp (SH)

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Chie Morimoto

Affiliation Department of Living Science Nutrition Course, Matsuyama Shinonome Junior College, Matsuyama, Ehime, Japan

Yoshimasa Inoue

Affiliation Carna Biosciences Incorporation, Kobe, Japan

Tsutomu Mizoshita

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Hironobu Tsukamoto

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Takaya Shimura

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Hiromi Kataoka

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Takeshi Kamiya

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Eiji Nishiwaki

Affiliation Carna Biosciences Incorporation, Kobe, Japan

Hiroshi Ishiguro

Affiliation Department of Medical Technology School of Health Sciences, Gifu University of Medical Science, Seki, Gifu, Japan

Shigeki Higashiyama

stanida@med.nagoya-cu.ac.jp (ST); shigeki@m.ehime-u.ac.jp (SH)

Affiliations Department of Cell Growth and Tumor Regulation, Proteo-Medicine Research Center, Ehime University, Toon, Ehime, Japan, Department of Biochemistry and Molecular Genetics, Ehime University Graduate School of Medicine, Ehime University, Toon, Ehime, Japan

Takashi Joh

Affiliation Department of Gastroenterology and Metabolism, Nagoya City University Graduate School of Medical Sciences, Nagoya, Aichi, Japan

Competing Interests

The authors have the following Competing interest: Yoshimasa Inoue and Eiji Nishiwaki are employed by commercial company Carna Biosciences Incorporation. There are no further patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors. All the authors declare that no financial conflict of interest exists in relation to the content of the article.

Author Contributions

Critical revision of the manuscript for important intellectual content: TJ SH. Statistical analysis: KO. Study supervision: ST TJ SH. Conceived and designed the experiments: ST TJ SH KO. Performed the experiments: KO CM. Analyzed the data: ST TJ SH TM HT TS HK TK KO. Contributed reagents/materials/analysis tools: YI EN HI SH. Wrote the paper: KO ST.