About the Authors

Naoko Isomura

Affiliation Department of Bioresources Engineering, Okinawa National College of Technology, Nago-City, Okinawa, Japan

Kenji Iwao

Affiliation Akajima Marine Science Laboratory, Zamamison, Okinawa, Japan

Hironobu Fukami

hirofukami@cc.miyazaki-u.ac.jp

Affiliation Department of Marine Biology and Environmental Science, Faculty of Agriculture, University of Miyazaki, Miyazaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Specimen collection: KI HF. Specimen identification: HF. Conceived and designed the experiments: NI HF. Performed the experiments: NI KI HF. Analyzed the data: NI HF. Contributed reagents/materials/analysis tools: NI KI HF. Wrote the paper: NI HF.