About the Authors

Qian Sheng

Contributed equally to this work with: Qian Sheng, Dingguo Cao

Affiliations Shandong Provincial Key Laboratory of Microbiological Engineering, Shandong Polytechnic University, Ji’nan, Shandong, China, Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China

Dingguo Cao

Contributed equally to this work with: Qian Sheng, Dingguo Cao

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Yan Zhou

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Qiuxia Lei

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Haixia Han

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Fuwei Li

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Yan Lu

cunfangwang@163.com (CW); luyan1957@163.com (YL)

Affiliations Institute of Poultry Science, Academy of Agricultural Sciences of Shandong Province, Ji’nan, Shandong, China, Poultry Breeding Engineering Technology Center of Shandong Province, Ji’nan, Shandong, China

Cunfang Wang

cunfangwang@163.com (CW); luyan1957@163.com (YL)

Affiliation Shandong Provincial Key Laboratory of Microbiological Engineering, Shandong Polytechnic University, Ji’nan, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CW YL DC. Performed the experiments: QS DC YZ QL. Analyzed the data: QS YZ. Contributed reagents/materials/analysis tools: YZ HH FL QL. Wrote the paper: QS CW.