About the Authors

Stefano Delli Pizzi

Affiliations ITAB, “G. D’Annunzio University”, Chieti, Italy, Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Cosmo Rossi

Affiliation Aging Research Center, Ce.S.I., “Gabriele d’Annunzio” University Foundation, Chieti, Italy

Vincenzo Di Matteo

Affiliation Laboratory of Neurophysiology, Istituto di Ricerche Farmacologiche Mario Negri, Consorzio Mario Negri Sud, Santa Maria Imbaro (Chieti), Italy

Ennio Esposito

Affiliation Laboratory of Neurophysiology, Istituto di Ricerche Farmacologiche Mario Negri, Consorzio Mario Negri Sud, Santa Maria Imbaro (Chieti), Italy

Simone Guarnieri

Affiliation Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Maria Addolorata Mariggiò

Affiliation Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Raffaella Franciotti

Affiliation ITAB, “G. D’Annunzio University”, Chieti, Italy

Massimo Caulo

Affiliations ITAB, “G. D’Annunzio University”, Chieti, Italy, Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Astrid Thomas

Affiliation Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Marco Onofrj

Affiliation Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Armando Tartaro

Contributed equally to this work with: Armando Tartaro, Laura Bonanni

Affiliations ITAB, “G. D’Annunzio University”, Chieti, Italy, Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Laura Bonanni

Contributed equally to this work with: Armando Tartaro, Laura Bonanni

l.bonanni@unich.it

Affiliation Department of Neuroscience and Imaging and CE.S.I. Aging Research Center, University G.d’Annunzio of Chieti-Pescara, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Gave substantial contributions to interpretation of data, revised the manuscript critically for important intellectual content and approved the version to be published: LB SDP CR VD EE RF SG MO A. Thomas A. Tartaro MAM MC. Conceived and designed the experiments: LB CR SG EE VD. Performed the experiments: CR SDP A. Thomas SG EE VD. Analyzed the data: SDP LB. Wrote the paper: LB SDP.