About the Authors

Jonathan Massey

jonathan.massey@manchester.ac.uk

Affiliation Centre for Integrated Genomic Medical Research (CIGMR), Institute of Population Health, Faculty of Medical and Human Sciences, The University of Manchester, Manchester, United Kingdom

Simon Rothwell

Affiliation Centre for Integrated Genomic Medical Research (CIGMR), Institute of Population Health, Faculty of Medical and Human Sciences, The University of Manchester, Manchester, United Kingdom

Clare Rusbridge

Affiliation Stone Lion Veterinary Hospital, London, United Kingdom

Anna Tauro

Affiliation Alcombe Veterinary Surgery, London, United Kingdom

Diane Addicott

Affiliation Hungarian Vizsla Breed Club, Royal Tunbridge Wells, United Kingdom

Hector Chinoy

Affiliation Rheumatic Diseases Centre, Manchester Academic Health Science Centre, The 6 University of Manchester, Salford Royal NHS Foundation Trust, Salford, United Kingdom

Robert G. Cooper

Affiliations Centre for Integrated Genomic Medical Research (CIGMR), Institute of Population Health, Faculty of Medical and Human Sciences, The University of Manchester, Manchester, United Kingdom, Rheumatic Diseases Centre, Manchester Academic Health Science Centre, The 6 University of Manchester, Salford Royal NHS Foundation Trust, Salford, United Kingdom

William E. R. Ollier

Affiliation Centre for Integrated Genomic Medical Research (CIGMR), Institute of Population Health, Faculty of Medical and Human Sciences, The University of Manchester, Manchester, United Kingdom

Lorna J. Kennedy

Affiliation Centre for Integrated Genomic Medical Research (CIGMR), Institute of Population Health, Faculty of Medical and Human Sciences, The University of Manchester, Manchester, United Kingdom

Competing Interests

Clare Rusbridge is affiliated with Stone Lion Veterinary Hospital and Anna Tauro with Alcombe Veterinary Surgery, both commercial companies. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Reviewed diagnosis: CR AT. Conceived and designed the experiments: JM SR CR HC RGC WERO LJK. Performed the experiments: JM SR. Analyzed the data: JM DA LJK. Contributed reagents/materials/analysis tools: CR AT DA. Wrote the paper: JM.