About the Authors

Bing He

Affiliation Division of Matrix Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, Stockholm, Sweden

Anne-May Österholm

Affiliation Division of Matrix Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, Stockholm, Sweden

Juha R. M. Ojala

Affiliation Division of Matrix Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, Stockholm, Sweden

Ann-Charlotte Andersson

Affiliation Division of Matrix Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, Stockholm, Sweden

Karl Tryggvason

karl.tryggvason@ki.se

Affiliation Division of Matrix Biology, Department of Medical Biochemistry and Biophysics, Karolinska Institute, Stockholm, Sweden

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BH. Performed the experiments: BH AMÖ JRO ACA. Analyzed the data: BH AMÖ JRO ACA. Contributed reagents/materials/analysis tools: KT. Wrote the paper: BH KT.