About the Authors

John Walmsley

Affiliation Department of Computer Science, University of Oxford, Oxford, United Kingdom

Jose F. Rodriguez

Affiliations Aragón Institute of Engineering Research, University of Zaragoza, Zaragoza, Spain, Biomedical Research Networking Center in Bioengineering, Biomaterials and Nanomedicine, Zaragoza, Spain

Gary R. Mirams

Affiliation Department of Computer Science, University of Oxford, Oxford, United Kingdom

Kevin Burrage

Affiliations Department of Computer Science, University of Oxford, Oxford, United Kingdom, School of Mathematical Sciences, Queensland University of Technology, Brisbane, Queensland, Australia

Igor R. Efimov

Affiliation Department of Biomedical Engineering, Washington University in St. Louis, St. Louis, Missouri, United States of America

Blanca Rodriguez

blanca@cs.ox.ac.uk

Affiliation Department of Computer Science, University of Oxford, Oxford, United Kingdom

Competing Interests

One of the authors on this paper, Dr. GM, has received research support from GlaxoSmithKline Ltd. The authors declare this as a competing interest. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Wrote simulation software: JW JR GM. Conceived and designed the experiments: JW KB IE BR. Performed the experiments: JW. Analyzed the data: JW JR GM BR. Contributed reagents/materials/analysis tools: IE. Wrote the paper: JW JR GM KB IE BR.