About the Authors

Charles J. Shoemaker

Affiliation Departmentof Cell Biology, University of Virginia, Charlottesville, Virginia, United States of America

Kathryn L. Schornberg

Affiliation Departmentof Cell Biology, University of Virginia, Charlottesville, Virginia, United States of America

Sue E. Delos

Affiliation Departmentof Cell Biology, University of Virginia, Charlottesville, Virginia, United States of America

Corinne Scully

Affiliation U.S. Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Hassan Pajouhesh

Affiliation Zalicus Inc., Cambridge, Massachusetts, United States of America

Gene G. Olinger

Affiliation U.S. Army Medical Research Institute of Infectious Diseases (USAMRIID), Fort Detrick, Maryland, United States of America

Lisa M. Johansen

Affiliation Zalicus Inc., Cambridge, Massachusetts, United States of America

Judith M. White

jw7g@virginia.edu

Affiliations Departmentof Cell Biology, University of Virginia, Charlottesville, Virginia, United States of America, Departmentof Microbiology, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

Lisa M. Johansen is and Hassan Pajouhesh was an employee of Zalicus, Inc. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CJS GGO LMJ JMW. Performed the experiments: CJS KLS SED CS. Analyzed the data: CJS KLS SED CS HP GGO JMW. Wrote the paper: CJS HP GGO LMJ JMW.