About the Authors

Le Wang

Affiliation Department of Veterinary and Animal Sciences, University of Massachusetts, Amherst, Massachusetts, United States of America

Erica A. Pawlak

Affiliation Department of Veterinary and Animal Sciences, University of Massachusetts, Amherst, Massachusetts, United States of America

Philip J. Johnson

Affiliation The University of Missouri College of Veterinary Medicine, Columbia, Missouri, United States of America

James K. Belknap

Affiliation The Ohio State University College of Veterinary Medicine, Columbus, Ohio, United States of America

Susan Eades

Affiliation The Louisiana State School of Veterinary Medicine, Baton Rouge, Louisiana, United States of America

Sharon Stack

Affiliation The University of Notre Dame, Notre Dame, Indiana, United States of America

Helene Cousin

Affiliation Department of Veterinary and Animal Sciences, University of Massachusetts, Amherst, Massachusetts, United States of America

Samuel J. Black

sblack@vasci.umass.edu

Affiliation Department of Veterinary and Animal Sciences, University of Massachusetts, Amherst, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LW EP HC SS SJB. Performed the experiments: LW EP PJJ JKB. Analyzed the data: LW EP HC SJB. Contributed reagents/materials/analysis tools: PJJ SE JKB. Wrote the paper: SJB LW EP.