About the Authors

Julia T. Vilstrup

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Andaine Seguin-Orlando

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Mathias Stiller

Affiliation Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Aurelien Ginolhac

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Maanasa Raghavan

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Sandra C. A. Nielsen

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Jacobo Weinstock

Affiliation Faculty of Humanities, University of Southampton, Southampton, United Kingdom

Duane Froese

Affiliation Department of Earth and Atmospheric Sciences, University of Alberta, Alberta, Canada

Sergei K. Vasiliev

Affiliation Institute of Archaeology and Ethnography, Russian Academy of Sciences, Novosibirsk, Russia

Nikolai D. Ovodov

Affiliation Laboratory of Archaeology and Paleogeography of Central Siberia, Institute of Archaeology and Ethnography, Russian Academy of Sciences, Novosibirsk, Russia

Joel Clary

Affiliation Centre de Conservation et d’Étude des Collections, Musée des Confluences, Lyon, France

Kristofer M. Helgen

Affiliation Division of Mammals, National Museum of Natural History, Smithsonian Institution, Washington D.C., United States of America

Robert C. Fleischer

Affiliation Center for Conservation and Evolutionary Genetics, Smithsonian National Zoological Park, Smithsonian Institution, Washington D.C., United States of America

Alan Cooper

Affiliation Australian Centre for Ancient DNA, School of Earth and Environmental Sciences, The University of Adelaide, South Australia, Australia

Beth Shapiro

Affiliation Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Ludovic Orlando

lorlando@snm.ku.dk

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Provided taxonomic and nomenclatural input: KMH. Conceived and designed the experiments: JTV LO MS BS. Performed the experiments: JTV ASO MS MR LO. Analyzed the data: JTV AG LO. Contributed reagents/materials/analysis tools: AC BS DF KMH JC JW RCF SCAN SKV NDO LO. Wrote the paper: JTV LO.