About the Authors

Naganori Kamiyama

Contributed equally to this work with: Naganori Kamiyama, Masahiro Yamamoto

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Masahiro Yamamoto

Contributed equally to this work with: Naganori Kamiyama, Masahiro Yamamoto

myamamoto@biken.osaka-u.ac.jp (MY); ktakeda@ongene.med.osaka-u.ac.jp (KT)

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Hiroyuki Saiga

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan

Ji Su Ma

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Jun Ohshima

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Sakaaki Machimura

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Miwa Sasai

Affiliations Laboratory of Immunoparasitology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Department of Immunoparasitology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Taishi Kimura

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan

Yoshiyasu Ueda

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan

Hisako Kayama

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Kiyoshi Takeda

myamamoto@biken.osaka-u.ac.jp (MY); ktakeda@ongene.med.osaka-u.ac.jp (KT)

Affiliations Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan, Laboratory of Mucosal Immunology, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NK MY KT. Performed the experiments: NK MY HS JSM SM TK. Analyzed the data: NK MY KT. Contributed reagents/materials/analysis tools: MS JO YU HK. Wrote the paper: NK MY KT.