About the Authors

Hélène Follet

Helene.Follet@inserm.fr

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Delphine Farlay

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Yohann Bala

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Stéphanie Viguet-Carrin

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Evelyne Gineyts

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Brigitte Burt-Pichat

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Julien Wegrzyn

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France, Département de Chirurgie Orthopédique, Pavillon T, Hôpital Edouard Herriot, Lyon, France

Pierre Delmas

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Georges Boivin

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Roland Chapurlat

Affiliations INSERM, UMR 1033, Lyon, France, Université de Lyon, UMR 1033, Lyon, France

Competing Interests

This study was supported in part by an unrestricted educational grant from Eli Lilly to INSERM. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: HF PD. Performed the experiments: HF DF YB SVC EG BBP. Analyzed the data: HF DF YB SVC EG. Contributed reagents/materials/analysis tools: HF JW PD. Wrote the paper: HF DF YB SVC GB RC.