About the Authors

Hyun-Ji Seo

Contributed equally to this work with: Hyun-Ji Seo, Heung Chul Kim

Affiliation Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea

Heung Chul Kim

Contributed equally to this work with: Hyun-Ji Seo, Heung Chul Kim

Affiliation 5th Medical Detachment, 168th Multifunctional Medical Battalion, 65th Medical Brigade, Unit 15247, Seoul, Republic of Korea

Terry A. Klein

Affiliation Force Health Protection and Preventive Medicine, 65th Medical Brigade/USAMEDDAC-Korea, Unit 15281, Seoul, Republic of Korea

Andrew M. Ramey

Affiliation US Geological Survey, Alaska Science Center, Anchorage, Arkansas, United States of America

Ji-Hye Lee

Affiliation Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea

Soon-Goo Kyung

Affiliation Korea Racing Authority, Gwacheon, Republic of Korea

Jee-Yong Park

Affiliation Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea

Yun Sang Cho

Affiliation Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea

In-Soo Cho

Affiliation Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea

Jung-Yong Yeh

malvab2848@gmail.com

Affiliations Foreign Animal Disease Division, Animal, Plant & Fisheries Quarantine & Inspection Agency, Anyang-si, Republic of Korea, Division of Life Sciences, College of Life Sciences and Bioengineering, University of Incheon, Incheon, Republic of Korea

Competing Interests

SGK are employed by Korea Racing Authority; AMR is employed by Alaska Science Center. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conducted field work: HJS HCK JHL JYY SGK JYP. Conceived and designed the experiments: JYY. Performed the experiments: HJS JHL HCK. Contributed reagents/materials/analysis tools: HCK TAK JYY YSC ISC AMR. Wrote the paper: JYY HJS HCK TAK AMR.