About the Authors

Detao Li

Contributed equally to this work with: Detao Li, Liwen Wang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Liwen Wang

Contributed equally to this work with: Detao Li, Liwen Wang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Xu Liu

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Dezhou Cui

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Tingting Chen

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Hua Zhang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Chuan Jiang

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Chunyan Xu

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Peng Li

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Song Li

Affiliation State Key Laboratory of Crop Biology, Shandong Agricultural University, Taian, Shandong, P. R. China

Li Zhao

hbchen@genetics.ac.cn (HC); zhaoli@genetics.ac.cn (LZ)

Affiliation State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, P. R. China

Huabang Chen

hbchen@genetics.ac.cn (HC); zhaoli@genetics.ac.cn (LZ)

Affiliation State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HC LW DL. Performed the experiments: DL LW. Analyzed the data: LW DL XL DC TC HZ CJ CX PL SL LZ. Wrote the paper: DL LW.