About the Authors

Li-Li Bai

Contributed equally to this work with: Li-Li Bai, Wei-Bo Yin

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Wei-Bo Yin

Contributed equally to this work with: Li-Li Bai, Wei-Bo Yin

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Yu-Hong Chen

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Li-Li Niu

Affiliation Laboratory of Proteomics, Institute of Biophysics, Chinese Academy of Sciences, Beijing, People’s Republic of China

Yong-Ru Sun

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Shi-Min Zhao

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Fu-Quan Yang

Affiliation Laboratory of Proteomics, Institute of Biophysics, Chinese Academy of Sciences, Beijing, People’s Republic of China

Richard R.-C. Wang

Affiliation USDA-ARS, FRRL, Utah State University, Logan, Utah, United States of America

Qing Wu

Affiliation Polo Biology Science Park Co., Ltd., Beijing, People’s Republic of China

Xiang-Qi Zhang

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Zan-Min Hu

zmhu@genetics.ac.cn

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Competing Interests

One author (Qing Wu), employed by a commercial company, Polo Biology Science Park Co., Ltd., Beijing, contributed to this paper in help drafting the manuscript. The authors declare the affiliation to this company and declare that this does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Reviewed and modified the manuscript: RRCW QW XQZ. Conceived and designed the experiments: YRS SMZ ZMH. Performed the experiments: LLB WBY YHC LLN. Analyzed the data: LLB WBY FQY ZMH. Contributed reagents/materials/analysis tools: LLB WBY YHC ZMH LLN FQY. Wrote the paper: LLB ZMH.