About the Authors

Yan Huang

Contributed equally to this work with: Yan Huang, Wenjun Chen

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Wenjun Chen

Contributed equally to this work with: Yan Huang, Wenjun Chen

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Xiaoyun Wang

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Hailiang Liu

Affiliation Guangzhou iGenomics Co., Ltd, Guangzhou, Guangdong, People’s Republic of China

Yangyi Chen

Affiliation Guangzhou iGenomics Co., Ltd, Guangzhou, Guangdong, People’s Republic of China

Lei Guo

Affiliation Guangzhou iGenomics Co., Ltd, Guangzhou, Guangdong, People’s Republic of China

Fang Luo

Affiliation Guangzhou iGenomics Co., Ltd, Guangzhou, Guangdong, People’s Republic of China

Jiufeng Sun

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Qiang Mao

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Pei Liang

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Zhizhi Xie

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Chenhui Zhou

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Yanli Tian

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Xiaoli Lv

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Lisi Huang

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Juanjuan Zhou

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Yue Hu

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Ran Li

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Fan Zhang

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Huali Lei

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Wenfang Li

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Xuchu Hu

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Chi Liang

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Jin Xu

yuxb@mail.sysu.edu.cn (XBY); xuerong2@mail.sysu.edu.cn (XRL); xujin@mail.sysu.edu.cn (JX)

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Xuerong Li

yuxb@mail.sysu.edu.cn (XBY); xuerong2@mail.sysu.edu.cn (XRL); xujin@mail.sysu.edu.cn (JX)

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Xinbing Yu

yuxb@mail.sysu.edu.cn (XBY); xuerong2@mail.sysu.edu.cn (XRL); xujin@mail.sysu.edu.cn (JX)

Affiliations Department of Parasitology, Zhongshan School of Medicine, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China, Key Laboratory for Tropical Diseases Control of Ministry of Education, Sun Yat-sen University, Guangzhou, Guangdong, People’s Republic of China

Competing Interests

HL. Liu, LG, YYC and FL employed by Guangzhou iGenomics Co., Ltd involved in helping the authors analyze the data. The authors have no other relationship with Guangzhou iGenomics Co., Ltd relating to employment, patents, products in development or marketed products etc. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: XBY XRL JX. Performed the experiments: YH WJC XYW FL. Analyzed the data: YH WJC HL. Liu LG YYC. Contributed reagents/materials/analysis tools: JFS QM PL CHZ YLT XLL LSH JJZ YH RL FZ HL. Lei WFL XCH CL. Wrote the paper: YH WJC HL. Liu LG YYC ZZX.