About the Authors

Naoki Kishimoto

Contributed equally to this work with: Naoki Kishimoto, Jun-ichi Nagai, Takehito Kinoshita

Affiliation Agrogenomics Research Center, National Institute of Agrobiological Sciences (NIAS), Tsukuba, Ibaraki, Japan

Jun-ichi Nagai

Contributed equally to this work with: Naoki Kishimoto, Jun-ichi Nagai, Takehito Kinoshita

Affiliation Tokunoshima Branch, Kagoshima Prefectural Institute for Agricultural Development (KIAD), Isen, Ōshima-gun, Kagoshima, Japan

Takehito Kinoshita

Contributed equally to this work with: Naoki Kishimoto, Jun-ichi Nagai, Takehito Kinoshita

Affiliation Vegetable Flower and Ornamental Plant Division, Saga Prefectural Agriculture Research Center, Kawasoe, Saga City, Saga, Japan

Keiichiro Ueno

Affiliation Kumage Branch, KIAD, Nishinoomote, Nishinoomote City, Kagoshima, Japan

Yuko Ohashi

Affiliation Division of Plant Sciences, NIAS, Tsukuba, Ibaraki, Japan

Ichiro Mitsuhara

mituhara@affrc.go.jp

Affiliation Division of Plant Sciences, NIAS, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IM. Performed the experiments: JN TK KU IM NK YO. Analyzed the data: IM NK. Wrote the paper: NK IM YO.