About the Authors

Koestan Gadan

Contributed equally to this work with: Koestan Gadan, Ane Sandtrø

Affiliation Norwegian School of Veterinary Science, Oslo, Norway

Ane Sandtrø

Contributed equally to this work with: Koestan Gadan, Ane Sandtrø

Affiliation Norwegian School of Veterinary Science, Oslo, Norway

Inderjit S. Marjara

Affiliation Norwegian School of Veterinary Science, Oslo, Norway

Nina Santi

Affiliation Aqua Gen AS, Trondheim, Norway

Hetron M. Munang'andu

Affiliation Norwegian School of Veterinary Science, Oslo, Norway

Øystein Evensen

oystein.evensen@nvh.no

Affiliation Norwegian School of Veterinary Science, Oslo, Norway

Competing Interests

Ane Sandtrø is currently affiliated with PHARMAQ which is a biologics company developing vaccines for aquaculture fish. The topics included in the paper has no direct economic implication to the work that Ane Sandtrø is currently doing. Nina Santi is employed with AquaGen AS which is a breeding company for Atlantic salmon. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NS OE. Performed the experiments: KG AS. Analyzed the data: KG AS ISM OE HMM. Contributed reagents/materials/analysis tools: KG AS IS NS HMM. Wrote the paper: OE KG ISM HMM.