About the Authors

Agnes E. Coutinho

Affiliations University/British Heart Foundation Centre for Cardiovascular Research, University of Edinburgh, Edinburgh, United Kingdom, Medical Research Council Centre for Inflammation Research, University of Edinburgh, Edinburgh, United Kingdom

Jeremy K. Brown

Affiliation Medical Research Council Centre for Reproductive Health, University of Edinburgh, Edinburgh, United Kingdom

Fu Yang

Affiliations University/British Heart Foundation Centre for Cardiovascular Research, University of Edinburgh, Edinburgh, United Kingdom, Medical Research Council Centre for Inflammation Research, University of Edinburgh, Edinburgh, United Kingdom

David G. Brownstein

Affiliation Mouse Pathology Core Laboratory, University of Edinburgh, Edinburgh, United Kingdom

Mohini Gray

Affiliation Medical Research Council Centre for Inflammation Research, University of Edinburgh, Edinburgh, United Kingdom

Jonathan R. Seckl

Affiliation University/British Heart Foundation Centre for Cardiovascular Research, University of Edinburgh, Edinburgh, United Kingdom

John S. Savill

Affiliation Medical Research Council Centre for Inflammation Research, University of Edinburgh, Edinburgh, United Kingdom

Karen E. Chapman

Karen.Chapman@ed.ac.uk

Affiliation University/British Heart Foundation Centre for Cardiovascular Research, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

JRS holds patents on the use of 11beta-HSD1 inhibitors in diabetes, atherosclerotic disease and age-associated cognitive impairment and has been pursuing the development of 11beta-HSD1 inhibitors in an academic setting with submitted patents comprising composition of matter. There are no further patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AEC MG JRS JSS KEC. Performed the experiments: AEC JKB FY. Analyzed the data: AEC DGG KEC. Contributed reagents/materials/analysis tools: AEC JKB FY MG. Wrote the paper: AEC KEC.