About the Authors

Øystein Brenna

Oystein.Brenna@stolav.no

Affiliations Department of Gastroenterology and Hepatology, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Marianne W. Furnes

Affiliation Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Ignat Drozdov

Affiliation Bering Limited, Richmond, United Kingdom

Atle van Beelen Granlund

Affiliation Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Arnar Flatberg

Affiliation Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Arne K. Sandvik

Affiliations Department of Gastroenterology and Hepatology, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Rosalie T. M. Zwiggelaar

Affiliation Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Ronald Mårvik

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Gastrointestinal Surgery, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway

Ivar S. Nordrum

Affiliations Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway, Department of Pathology and Medical Genetics, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway

Mark Kidd

Affiliation Department of Surgery, Section of Gastroenterology, Yale School of Medicine, New Haven, Connecticut, United States of America

Björn I. Gustafsson

Affiliations Department of Gastroenterology and Hepatology, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway, Department of Cancer Research and Molecular Medicine, Norwegian University of Science and Technology, Trondheim, Norway

Competing Interests

The authors have the following interests. Ignat Drozdov is employed by Bering Limited and was paid was paid by NTNU as a consultant to do this work. Bering Limited was involved in data analysis. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BG ØB MWF RM. Performed the experiments: ØB MWF BG AvBG RTMZ. Analyzed the data: ØB BG ID AvBG AF AS ISN MK. Contributed reagents/materials/analysis tools: ØB BG ID AvBG AF AS RM. Wrote the paper: ØB MWF ID AvBG AS ISN MK BG.