About the Authors

Masaki Yoda

Affiliations Anti-aging Orthopedic Research, School of Medicine, Keio University, Tokyo, Japan, Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Tokuhiro Kimura

Affiliation Pathology, School of Medicine, Keio University, Tokyo, Japan

Takahide Tohmonda

Affiliations Anti-aging Orthopedic Research, School of Medicine, Keio University, Tokyo, Japan, Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Hideo Morioka

Affiliation Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Morio Matsumoto

Affiliation Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Yasunori Okada

Affiliation Pathology, School of Medicine, Keio University, Tokyo, Japan

Yoshiaki Toyama

Affiliation Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Keisuke Horiuchi

horiuchi@z3.keio.jp

Affiliations Anti-aging Orthopedic Research, School of Medicine, Keio University, Tokyo, Japan, Department of Orthopedic Surgery, School of Medicine, Keio University, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed histological analysis: TK YO. Conceived and designed the experiments: KH. Performed the experiments: MY KH. Analyzed the data: MY TT HM MM YT KH. Wrote the paper: KH.