About the Authors

Naofumi Takahashi

Affiliations AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Takushi Nomura

Affiliations AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Yusuke Takahara

Affiliations AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Hiroyuki Yamamoto

Affiliation AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan

Teiichiro Shiino

Affiliation AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan

Akiko Takeda

Affiliation AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan

Makoto Inoue

Affiliation DNAVEC Corporation, Tsukuba, Japan

Akihiro Iida

Affiliation DNAVEC Corporation, Tsukuba, Japan

Hiroto Hara

Affiliation DNAVEC Corporation, Tsukuba, Japan

Tsugumine Shu

Affiliation DNAVEC Corporation, Tsukuba, Japan

Mamoru Hasegawa

Affiliation DNAVEC Corporation, Tsukuba, Japan

Hiromi Sakawaki

Affiliation Institute for Virus Research, Kyoto University, Kyoto, Japan

Tomoyuki Miura

Affiliation Institute for Virus Research, Kyoto University, Kyoto, Japan

Tatsuhiko Igarashi

Affiliation Institute for Virus Research, Kyoto University, Kyoto, Japan

Yoshio Koyanagi

Affiliation Institute for Virus Research, Kyoto University, Kyoto, Japan

Taeko K. Naruse

Affiliation Department of Molecular Pathogenesis, Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan

Akinori Kimura

Affiliation Department of Molecular Pathogenesis, Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan

Tetsuro Matano

tmatano@nih.go.jp

Affiliations AIDS Research Center, National Institute of Infectious Diseases, Tokyo, Japan, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Competing Interests

Makoto Inoue, Akihiro Iida, Hiroto Hara, Tsugumine Shu and Mamoru Hasegawa are employed by DNAVEC Corporation. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Performed animal experiments: HS TM TI YK. Performed MHC-I typing: TKN AK. Conceived and designed the experiments: NT TM. Performed the experiments: NT TN YT HY AT. Analyzed the data: NT HY T. Shiino TM. Contributed reagents/materials/analysis tools: MI AI HH T. Shu MH. Wrote the paper: NT TM.