About the Authors

Laixiang Xu

xulx@mail.qfnu.edu.cn

Affiliation College of Life Sciences, Qufu Normal University, Qufu, Shandong, People’s Republic of China

Huiliang Xue

Affiliation College of Life Sciences, Qufu Normal University, Qufu, Shandong, People’s Republic of China

Mingjing Song

Affiliations Institute of Laboratory Animal Science, Chinese Academy of Medical Sciences(CAMS) & Comparative Medicine Centre, Peking Union Medical College (PUMC), Beijing, People’s Republic of China, State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Qinghua Zhao

Affiliation College of Life Sciences, Qufu Normal University, Qufu, Shandong, People’s Republic of China

Jingping Dong

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Juan Liu

Affiliation College of Life Sciences, Qufu Normal University, Qufu, Shandong, People’s Republic of China

Yu Guo

Affiliation College of Life Sciences, Qufu Normal University, Qufu, Shandong, People’s Republic of China

Tongqin Xu

Affiliation Wugong Station for Pest Monitoring and Forecasting, Raoyang County, Hebei Province, People’s Republic of China

Xiaoping Cao

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Fusheng Wang

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Shuqing Wang

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Shushen Hao

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Hefang Yang

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Zhibin Zhang

Affiliation State Key Laboratory of Integrated Management on Pest Insects and Rodents in Agriculture, Institute of Zoology, Chinese Academy of Sciences, Beijing, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Contributed reagents: QZ JL XC. Contributed materials: TX FW SW SH HY. Contributed analysis tools: JD YG LX. Conceived and designed the experiments: LX ZZ. Performed the experiments: LX HX. Analyzed the data: HX LX MS ZZ. Wrote the paper: HX LX.