About the Authors

Shin Wakimoto

Affiliation Department of Immunology and Parasitology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Haruyuki Nakayama-Imaohji

Affiliation Department of Microbiology, Faculty of Medicine, Kagawa University, Kagawa, Japan

Minoru Ichimura

Affiliations Department of Immunology and Parasitology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan, Department of Microbiology, Faculty of Medicine, Kagawa University, Kagawa, Japan

Hidetoshi Morita

Affiliation School of Veterinary Medicine, Azabu University, Kanagawa, Japan

Hideki Hirakawa

Affiliation Kazusa DNA Research Institute, Chiba, Japan

Tetsuya Hayashi

Affiliation Frontier Science Research Center, University of Miyazaki, Miyazaki, Japan

Koji Yasutomo

Affiliation Department of Immunology and Parasitology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Tomomi Kuwahara

tomomi@med.kagawa-u.ac.jp

Affiliation Department of Microbiology, Faculty of Medicine, Kagawa University, Kagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SW HNI TH KY TK. Performed the experiments: SW HNI MI TK. Analyzed the data: SW HNI HH TH TK. Contributed reagents/materials/analysis tools: HH HM. Wrote the paper: SW TK.