About the Authors

Yepeng Sun

Affiliations Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China, College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Fawei Wang

Affiliation Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China

Nan Wang

Affiliation Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China

Yuanyuan Dong

Affiliation Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China

Qi Liu

Affiliation Institute of Genomic Medicine, Wenzhou Medical College, Wenzhou, China

Lei Zhao

Affiliation Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species (GBOWS), Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China

Huan Chen

Affiliations Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China, College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Weican Liu

Affiliations Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China, College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Hailong Yin

Affiliations Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China, College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Xiaomei Zhang

Affiliation College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Yanxi Yuan

Affiliation College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Haiyan Li

hyli99@163.com

Affiliations Ministry of Education Engineering Research Center of Bioreactor and Pharmaceutical Development, Jilin Agricultural University, Changchun, Jilin, China, College of Life Sciences, Jilin Agricultural University, Changchun, Jilin, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Read and approved the final manuscript: YPS FWW NW YYD QL LZ HC WL HLY XZ YXY HYL. Conceived and designed the experiments: HYL YPS. Performed the experiments: YPS YXY HLY WL. Analyzed the data: YPS QL LZ. Contributed reagents/materials/analysis tools: YPS QL LZ. Wrote the paper: HYL YPS YYD FWW.