About the Authors

Seungbok Lee

Contributed equally to this work with: Seungbok Lee, Seung Hwan Paik

Affiliations Genomic Medicine Institute (GMI), Medical Research Center, Seoul National University, Seoul, Korea, Department of Biomedical Sciences, Seoul National University Graduate School, Seoul, Korea

Seung Hwan Paik

Contributed equally to this work with: Seungbok Lee, Seung Hwan Paik

Affiliation Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea

Hyun-Jin Kim

Affiliations Genomic Medicine Institute (GMI), Medical Research Center, Seoul National University, Seoul, Korea, Department of Biomedical Sciences, Seoul National University Graduate School, Seoul, Korea

Hyeong Ho Ryu

Affiliation Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea

Soeun Cha

Affiliation Genomic Medicine Institute (GMI), Medical Research Center, Seoul National University, Seoul, Korea

Seong Jin Jo

Affiliation Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea

Hee Chul Eun

Affiliations Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea, Laboratory of Cutaneous Aging and Hair Research, Clinical Research Institute, Seoul National University Hospital, Seoul, Korea, Institute of Dermatological Science, Seoul National University College of Medicine, Seoul, Korea

Jeong-Sun Seo

Affiliations Genomic Medicine Institute (GMI), Medical Research Center, Seoul National University, Seoul, Korea, Department of Biomedical Sciences, Seoul National University Graduate School, Seoul, Korea, Department of Biochemistry and Molecular Biology, Seoul National University College of Medicine, Seoul, Korea, Psoma Therapeutics Inc., Seoul, Korea, Macrogen Inc., Seoul, Korea

Jong-Il Kim

jongil@snu.ac.kr (JK); oskwon@snu.ac.kr (OSK)

Affiliations Genomic Medicine Institute (GMI), Medical Research Center, Seoul National University, Seoul, Korea, Department of Biomedical Sciences, Seoul National University Graduate School, Seoul, Korea, Department of Biochemistry and Molecular Biology, Seoul National University College of Medicine, Seoul, Korea, Psoma Therapeutics Inc., Seoul, Korea

Oh Sang Kwon

jongil@snu.ac.kr (JK); oskwon@snu.ac.kr (OSK)

Affiliations Department of Dermatology, Seoul National University College of Medicine, Seoul, Korea, Laboratory of Cutaneous Aging and Hair Research, Clinical Research Institute, Seoul National University Hospital, Seoul, Korea, Institute of Dermatological Science, Seoul National University College of Medicine, Seoul, Korea

Competing Interests

JS is a PLOS ONE Editorial Board member. JS is a consultant for Macrogen and Psoma Therapeutics, and JK is a consultant for Psoma Therapeutics. This does not alter the authors' adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JK OSK. Performed the experiments: SL SC. Analyzed the data: SL SHP HK. Contributed reagents/materials/analysis tools: HCE JS JK OSK. Wrote the paper: SL SHP HHR SJJ.