About the Authors

Tetsufumi Takahashi

Affiliations Division of Gastroenterology and Hepatology, Department of Internal Medicine, School of Medicine, Keio University, Tokyo, Japan, Laboratory of Pathophysiology, Division of Clinical Medicine, School of Pharmacy, Kitasato University, Tokyo, Japan

Hidekazu Suzuki

hsuzuki@a6.keio.jp

Affiliation Division of Gastroenterology and Hepatology, Department of Internal Medicine, School of Medicine, Keio University, Tokyo, Japan

Takao Imai

Affiliation Department of Physiology, School of Medicine, Keio University, Tokyo, Japan

Shinsuke Shibata

Affiliation Department of Physiology, School of Medicine, Keio University, Tokyo, Japan

Yoshiaki Tabuchi

Affiliation Division of Molecular Genetics, Life Science Research Center, University of Toyama, Toyama, Japan

Kanji Tsuchimoto

Affiliation Laboratory of Pathophysiology, Division of Clinical Medicine, School of Pharmacy, Kitasato University, Tokyo, Japan

Hideyuki Okano

Affiliation Department of Physiology, School of Medicine, Keio University, Tokyo, Japan

Toshifumi Hibi

Affiliation Division of Gastroenterology and Hepatology, Department of Internal Medicine, School of Medicine, Keio University, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TT HS TI HO. Performed the experiments: TT. Analyzed the data: TT HS TI SS HO. Contributed reagents/materials/analysis tools: SS YT HO. Wrote the paper: TT HS TI SS HO. Supervised the experiments and their analysis: KT HO TH.