About the Authors

Jonathan Carlo Briones

Contributed equally to this work with: Jonathan Carlo Briones, Cheng-Han Tsai

Affiliation Graduate School, University of Santo Tomas, Manila, Philippines

Cheng-Han Tsai

Contributed equally to this work with: Jonathan Carlo Briones, Cheng-Han Tsai

Affiliation Institute of Oceanography, National Taiwan University, Taipei, Taiwan

Takefumi Nakazawa

take.nkzw@gmail.com

Affiliations Institute of Oceanography, National Taiwan University, Taipei, Taiwan, Center for Ecological Research, Kyoto University, Otsu, Japan

Yoichiro Sakai

Affiliation Center for Ecological Research, Kyoto University, Otsu, Japan

Rey Donne S. Papa

Affiliation Department of Biological Sciences and Research Center for the Natural and Applied Sciences, University of Santo Tomas, Manila, Philippines

Chih-hao Hsieh

Affiliations Institute of Oceanography, National Taiwan University, Taipei, Taiwan, Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei, Taiwan

Noboru Okuda

Affiliation Center for Ecological Research, Kyoto University, Otsu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RDP CHH. Performed the experiments: JCB CHT. Analyzed the data: JCB CHT. Contributed reagents/materials/analysis tools: NO YS. Wrote the paper: TN.