About the Authors

Yoshinobu Uno

Affiliation Department of Applied Molecular Biosciences, Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, Japan

Chizuko Nishida

Affiliation Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, Japan

Hiroshi Tarui

Current address: Omics Science Center, Yokohama Institute, RIKEN, Yokohama, Japan

Affiliation Genome Resource and Analysis Subunit, RIKEN Center for Developmental Biology, Kobe, Japan

Satoshi Ishishita

Affiliation Department of Applied Molecular Biosciences, Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, Japan

Chiyo Takagi

Affiliation Division of Morphogenesis, National Institute for Basic Biology, Okazaki, Japan

Osamu Nishimura

Affiliations Genome Resource and Analysis Subunit, RIKEN Center for Developmental Biology, Kobe, Japan, Department of Biophysics, Graduate School of Science, Kyoto University, Kyoto, Japan, Global COE Program for Evolution and Biodiversity, Graduate School of Science, Kyoto University, Kyoto, Japan

Junko Ishijima

Affiliation Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, Japan

Hidetoshi Ota

Affiliation Institute of Natural and Environmental Sciences and Museum of Nature and Human Activities, University of Hyogo, Hyogo, Japan

Ayumi Kosaka

Affiliation Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, Japan

Kazumi Matsubara

Current address: Wildlife Genetics Laboratory, Institute for Applied Ecology, University of Canberra, ACT, Australia

Affiliation Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, Japan

Yasunori Murakami

Current address: Graduate school of Science and Engineering, Ehime University, Matsuyama, Japan

Affiliation Laboratory for Evolutionary Morphology, RIKEN Center for Developmental Biology, Kobe, Japan

Shigeru Kuratani

Affiliation Laboratory for Evolutionary Morphology, RIKEN Center for Developmental Biology, Kobe, Japan

Naoto Ueno

Affiliations Division of Morphogenesis, National Institute for Basic Biology, Okazaki, Japan, Department of Basic Biology, School of Life Science, The Graduate University of Advanced Studies (SOKENDAI), Okazaki, Japan

Kiyokazu Agata

Affiliation Department of Biophysics, Graduate School of Science, Kyoto University, Kyoto, Japan

Yoichi Matsuda

yoimatsu@agr.nagoya-u.ac.jp

Affiliation Department of Applied Molecular Biosciences, Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Y. Matsuda YU KA. Performed the experiments: YU Y. Matsuda CN HT ON CT JI AK KM. Analyzed the data: YU HT CN SI Y. Matsuda. Contributed reagents/materials/analysis tools: HT ON CT Y. Murakami HO SK NU KA. Wrote the paper: YU Y. Matsuda.