About the Authors

Juan Martin Caballero

Contributed equally to this work with: Juan Martin Caballero, Ana Garzón

Affiliation Laboratory Animal Unit, Barcelona Biomedical Research Park, Barcelona, Spain

Ana Garzón

Contributed equally to this work with: Juan Martin Caballero, Ana Garzón

Current address: Azurebio S.L., Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Leticia González-Cintado

Affiliation Department of Immunology and Oncology, Centro Nacional de Biotecnología/CSIC, Madrid, Spain

Wioleta Kowalczyk

Current address: CSIRO Materials Science and Engineering, Australia

Affiliation Department of Experimental and Health Sciences, Pompeu Fabra University, Barcelona Biomedical Research Park, Barcelona, Spain

Ignacio Jimenez Torres

Current address: Igenbiotech, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Gloria Calderita

Current address: Nimgenetics, Parque Científico de Madrid, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Margarita Rodriguez

Current address: Hospital Universitario Sanchinarro, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Virgínia Gondar

Current address: Hospital Universitario de la Princesa, Unidad de Biología Molecular, Instituto Investigación Sanitaria Princesa, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Juan Jose Bernal

Current address: Protein Alternatives S.L., Tres Cantos, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Carlos Ardavín

Affiliation Department of Immunology and Oncology, Centro Nacional de Biotecnología/CSIC, Madrid, Spain

David Andreu

Affiliation Department of Experimental and Health Sciences, Pompeu Fabra University, Barcelona Biomedical Research Park, Barcelona, Spain

Thomas Zürcher

Current address: ForMune S.L., Vivero de Innovación, Navarra, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Cayetano von Kobbe

cvonkobbe@cbm.uam.es

Current address: Molecular Biology Centre “Severo Ochoa”, Spanish Research Council (CBMSO-CSIC), Universidad Autónoma de Madrid, Madrid, Spain

Affiliation Cancer Vaccines Unit, R & D Department, Chimera Pharma S.L.U., Madrid, Spain

Competing Interests

AG, IJT, GC, MR, VG, JJB, TZ and CvK were employed by Chimera Pharma S.L.U. during the course of this project. Currently Chimera Pharma is under judicial liquidation after a meeting of creditorś period of time. None of the authors belongs to Chimera anymore. The VLP-E7 and VLP-E7-B constructs described in this paper are now propietary of VLP The Vaccineś Company S.L. (VLP Bio) and protected under PCT WO 2011/054996. The new owners of the intellectual property rights agree with the publication of these data. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CvK TZ JMC CA. Performed the experiments: JMC AG LGC GC MR VG. Analyzed the data: JMC AG CvK TZ DA CA. Contributed reagents/materials/analysis tools: JJB DA IJT WK. Wrote the paper: CvK.