About the Authors

Sakiko Masuda

Affiliation Graduate School of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Sari Iwasaki

Affiliations Faculty of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan, Department of Pathology, Hokkaido University Graduate School of Medicine, Kita-ku, Sapporo, Japan

Utano Tomaru

Affiliation Department of Pathology, Hokkaido University Graduate School of Medicine, Kita-ku, Sapporo, Japan

Juri Sato

Affiliation School of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Ai Kawakami

Affiliation Graduate School of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Kana Ichijo

Affiliation School of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Sayuri Sogo

Affiliation School of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Tomohisa Baba

Current address: Division of Molecular Bioregulation, Cancer Research Institute, Kanazawa University, Kanazawa, Japan

Affiliation Department of Pathology, Hokkaido University Graduate School of Medicine, Kita-ku, Sapporo, Japan

Kazuaki Katsumata

Affiliation Faculty of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Masanori Kasahara

Affiliation Department of Pathology, Hokkaido University Graduate School of Medicine, Kita-ku, Sapporo, Japan

Akihiro Ishizu

aishizu@med.hokudai.ac.jp

Affiliation Faculty of Health Sciences, Hokkaido University, Kita-ku, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: UT AI. Performed the experiments: SM SI JS AK KI SS. Analyzed the data: SM SI UT JS AK KI SS TB KK AI. Contributed reagents/materials/analysis tools: UT MK AI. Wrote the paper: UT AI.