About the Authors

Akemi Shodai

Contributed equally to this work with: Akemi Shodai, Akemi Ido

Affiliation Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Akemi Ido

Contributed equally to this work with: Akemi Shodai, Akemi Ido

Affiliation Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Noriko Fujiwara

Affiliation Department of Biochemistry, Hyogo College of Medicine, Hyogo, Japan

Takashi Ayaki

Affiliation Department of Neurology, Kyoto University, Kyoto, Japan

Toshifumi Morimura

Affiliation Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Miki Oono

Affiliations Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan, Department of Neurology, Kyoto University, Kyoto, Japan

Tsukasa Uchida

Affiliations Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan, Department of Neurology, Kyoto University, Kyoto, Japan

Ryosuke Takahashi

Affiliation Department of Neurology, Kyoto University, Kyoto, Japan

Hidefumi Ito

Affiliations Department of Neurology, Kyoto University, Kyoto, Japan, Department of Neurology, Wakayama Medical University Graduate School of Medicine, Wakayama, Japan

Makoto Urushitani

uru@belle.shiga-med.ac.jp

Affiliation Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NF HI MU. Performed the experiments: AS AI TA MO TU. Analyzed the data: AI NF TM RT HI MU. Contributed reagents/materials/analysis tools: TA HI. Wrote the paper: AS NF TA HI MU.