About the Authors

Kristin M. Narayan

Affiliations Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America, Altravax, Inc., Sunnyvale, California, United States of America

Nitish Agrawal

Affiliation Department of Immunology and Microbial Sciences, The Scripps Research Institute, La Jolla, California, United States of America

Sean X. Du

Current address: Simcere Pharmaceutical Group, Institute of Vaccine Research, Nanjing, People's Republic of China

Affiliations Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America, Altravax, Inc., Sunnyvale, California, United States of America

Janelle E. Muranaka

Affiliations Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America, Altravax, Inc., Sunnyvale, California, United States of America

Katherine Bauer

Affiliation Department of Immunology and Microbial Sciences, The Scripps Research Institute, La Jolla, California, United States of America

Daniel P. Leaman

Affiliation Department of Immunology and Microbial Sciences, The Scripps Research Institute, La Jolla, California, United States of America

Pham Phung

Affiliation Monogram Biosciences, Inc., San Francisco, California, United States of America

Kay Limoli

Affiliation Monogram Biosciences, Inc., San Francisco, California, United States of America

Helen Chen

Affiliation Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America

Rebecca I. Boenig

Affiliations Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America, Altravax, Inc., Sunnyvale, California, United States of America

Terri Wrin

Affiliation Monogram Biosciences, Inc., San Francisco, California, United States of America

Michael B. Zwick

zwick@scripps.edu (MBZ); robert.whalen@altravax.com (RGW)

Affiliation Department of Immunology and Microbial Sciences, The Scripps Research Institute, La Jolla, California, United States of America

Robert G. Whalen

zwick@scripps.edu (MBZ); robert.whalen@altravax.com (RGW)

Affiliations Department of Infectious Diseases, Maxygen, Inc., Redwood City, California, United States of America, Altravax, Inc., Sunnyvale, California, United States of America

Competing Interests

KMN, SXD, JEM, HC, RIB, and RGW were employees of Maxygen, Inc. and/or Altravax, Inc. during this work. PP and TW are employees of Monogram Biosciences. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KMN NA SXD MBZ RGW. Performed the experiments: KMN NA JEM KB DPL PP KL HC RIB. Analyzed the data: KMN NA SXD PP TW MBZ RGW. Wrote the paper: KMN NA SXD TW MBZ RGW.