About the Authors

Wenming Jiang

Contributed equally to this work with: Wenming Jiang, Shuo Liu

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Shuo Liu

Contributed equally to this work with: Wenming Jiang, Shuo Liu

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Guangyu Hou

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Jinping Li

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Qingye Zhuang

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Suchun Wang

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Peng Zhang

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Jiming Chen

jmchen66@yahoo.cn

Affiliation The Laboratory of Avian Disease Surveillance, China Animal Health and Epidemiology Center, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JC WJ SL. Performed the experiments: WJ SL GH JL QZ SW PZ JC. Analyzed the data: JC SL WJ. Contributed reagents/materials/analysis tools: JC. Wrote the paper: JC WJ SL.