About the Authors

Joji Tsunada

tsunada@mail.med.upenn.edu

Current address: Department of Otorhinolaryngology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Affiliations Laboratory of Cognitive Neurobiology, Hokkaido University Graduate School of Medicine, Sapporo, Hokkaido, Japan, Humanity Neuroscience Institute, Sapporo, Hokkaido, Japan

Toshiyuki Sawaguchi

Affiliations Laboratory of Cognitive Neurobiology, Hokkaido University Graduate School of Medicine, Sapporo, Hokkaido, Japan, Humanity Neuroscience Institute, Sapporo, Hokkaido, Japan, Musashino Gakuin University, Sayama, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JT TS. Performed the experiments: JT. Analyzed the data: JT. Wrote the paper: JT TS.