About the Authors

Xuhao Pan

Contributed equally to this work with: Xuhao Pan, Qijun Zhang

Affiliations Rice Research Institute, Sichuan Agricultural University, Chengdu, China, Rice Research and Extension Center, University of Arkansas, Stuttgart, Arkansas, United States of America, USDA-ARS, Dale Bumpers National Rice Research Center, Stuttgart, Arkansas, United States of America

Qijun Zhang

Contributed equally to this work with: Xuhao Pan, Qijun Zhang

Affiliation Institute of Food Crops, Jiangsu Academy of Agricultural Sciences, Nanjing, China

Wengui Yan

Wengui.Yan@ars.usda.gov (WY); lishigui_sc@263.net (SL)

Affiliation USDA-ARS, Dale Bumpers National Rice Research Center, Stuttgart, Arkansas, United States of America

Melissa Jia

Affiliation USDA-ARS, Dale Bumpers National Rice Research Center, Stuttgart, Arkansas, United States of America

Aaron Jackson

Affiliation USDA-ARS, Dale Bumpers National Rice Research Center, Stuttgart, Arkansas, United States of America

Xiaobai Li

Affiliation State Key Lab of Rice Biology, Institute of Nuclear-Agriculture Sciences, Zhejiang University, Hangzhou, China

Limeng Jia

Affiliation The Institute of Virology and Biotechnology, Zhejiang Academy of Agriculture Sciences, Zhejiang, China

Bihu Huang

Affiliation University of Arkansas at Pine Bluff, Pine Bluff, Arkansas, United States of America

Peizhou Xu

Affiliation Rice Research Institute, Sichuan Agricultural University, Chengdu, China

Fernando Correa-Victoria

Affiliation RiceTec, Inc., Alvin, Texas, United States of America

Shigui Li

Wengui.Yan@ars.usda.gov (WY); lishigui_sc@263.net (SL)

Affiliation USDA-ARS, Dale Bumpers National Rice Research Center, Stuttgart, Arkansas, United States of America

Competing Interests

The authors declare that no conflict of interest exists in the submission of this manuscript that is approved by all authors for publication, and there are no competing interests from the affiliations of the authors including RiceTec., Inc. The authors further declare the affiliation(s) to this company (Fernando Correa-Victoria), along with any other relevant declarations relating to employment, consultancy, patents, products in development or marketed products etc. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: WY FC SL. Analyzed the data: XP QZ MJ AJ XL LJ BH PX. Contributed reagents/materials/analysis tools: WY MJ AJ FC. Wrote the paper: XP WY QZ AJ.