About the Authors

Fengjun Sun

Contributed equally to this work with: Fengjun Sun, He Gao, Yiquan Zhang

Affiliations State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China, Department of Pharmacy, Southwest Hospital, the Third Military Medical University, Chongqing, China

He Gao

Contributed equally to this work with: Fengjun Sun, He Gao, Yiquan Zhang

Affiliations State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China, State Key Laboratory for Infectious Disease Prevention and Control, National Institute for Communicable Disease Control and Prevention, Chinese Centre for Disease Control and Prevention, Beijing, China

Yiquan Zhang

Contributed equally to this work with: Fengjun Sun, He Gao, Yiquan Zhang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Li Wang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Nan Fang

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Yafang Tan

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Zhaobiao Guo

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Peiyuan Xia

py_xia@yahoo.com.cn (PX); dongshengzhou1977@gmail.com (DZ); ruifuyang@gmail.com (RY)

Affiliation Department of Pharmacy, Southwest Hospital, the Third Military Medical University, Chongqing, China

Dongsheng Zhou

py_xia@yahoo.com.cn (PX); dongshengzhou1977@gmail.com (DZ); ruifuyang@gmail.com (RY)

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Ruifu Yang

py_xia@yahoo.com.cn (PX); dongshengzhou1977@gmail.com (DZ); ruifuyang@gmail.com (RY)

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Competing Interests

Co-author Dongsheng Zhou is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DZ RY. Performed the experiments: FS HG YZ LW NF YT ZG. Analyzed the data: DZ FS PX. Contributed reagents/materials/analysis tools: DZ FS PX. Wrote the paper: DZ FS RY.