About the Authors

Ho Bin Jang

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Young Kyu Kim

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Carmelo S. del Castillo

Current address: Marine Animal Diseases Laboratory, School of Marine and Atmospheric Sciences, Stony Brook University, Stony Brook, New York, United States of America

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Seong Won Nho

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

In Seok Cha

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Seong Bin Park

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Mi Ae Ha

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Jun-ichi Hikima

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Sung Jong Hong

Affiliation Department of Medical Environmental Biology, College of Medicine, Chung-Ang University, DongJak-Gu, Seoul, South Korea

Takashi Aoki

Affiliations Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea, Consolidated Research Institute for Advanced Science and Medical Care, Waseda University, Shinjuku-ku, Tokyo, Japan

Tae Sung Jung

jungts@gnu.ac.kr

Affiliation Aquatic Biotechnology Center of WCU Project, College of Veterinary Medicine, Gyeongsang National University, Jinju, South Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HBJ TSJ TA SJH. Performed the experiments: HBJ YKK MAH. Analyzed the data: HBJ. Contributed reagents/materials/analysis tools: CSC SWN ISC SBP JH. Wrote the paper: HBJ.