About the Authors

Wentao Song

Contributed equally to this work with: Wentao Song, Yangzhen Li, Yongwei Zhao

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, Weihai Vocational College, Department of Biological and Chemical Engineering, Weihai, China

Yangzhen Li

Contributed equally to this work with: Wentao Song, Yangzhen Li, Yongwei Zhao

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China

Yongwei Zhao

Contributed equally to this work with: Wentao Song, Yangzhen Li, Yongwei Zhao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Yang Liu

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China

Yuze Niu

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China

Renyi Pang

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Guidong Miao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Xiaolin Liao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Changwei Shao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Fengtao Gao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Songlin Chen

chensl@ysfri.ac.cn

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SC. Performed the experiments: WS Y. Li YZ Y. Liu. Analyzed the data: WS YZ. Contributed reagents/materials/analysis tools: YN RP GM XL CS FG. Wrote the paper: WS SC. Revised the manuscript: SC.