About the Authors

Munekazu Naito

Contributed equally to this work with: Munekazu Naito, Shuichi Hirai

Affiliation Department of Anatomy, Tokyo Medical University, Tokyo, Japan

Shuichi Hirai

Contributed equally to this work with: Munekazu Naito, Shuichi Hirai

shinamon611@gmail.com

Affiliation Department of Anatomy, Tokyo Medical University, Tokyo, Japan

Makoto Mihara

Affiliation Department of Plastic and Reconstructive Surgery, Graduate School of Medicine, University of Tokyo, Tokyo, Japan

Hayato Terayama

Affiliation Department of Anatomy, Tokyo Medical University, Tokyo, Japan

Naoyuki Hatayama

Affiliation Department of Anatomy, Tokyo Medical University, Tokyo, Japan

Shogo Hayashi

Affiliation Medicinal Education Center, School of Medicine, Aichi Medical University, Aichi, Japan

Masayuki Matsushita

Affiliation Molecular and Cellular Physiology, University of the Ryukyus, Okinawa, Japan

Masahiro Itoh

Affiliation Department of Anatomy, Tokyo Medical University, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M. Naito S. Hirai M. Matsushita. Performed the experiments: M. Naito S. Hirai HT M. Matsushita. Analyzed the data: M. Naito S. Hirai HT M. Matsushita M. Mihara NH S. Hayashi MI. Contributed reagents/materials/analysis tools: MN S. Hirai S. Hayashi M. Matsushita MI. Wrote the paper: MN S. Hirai. Discussed the results and commented on the manuscript: MN S. Hirai M. Mihara HT NH S. Hayashi M. Matsushita MI.