About the Authors

Michael T. Cheeseman

m.cheeseman@har.mrc.ac.uk

Affiliations Medical Research Council Mammalian Genetics Unit, Medical Research Council Harwell, Oxfordshire, United Kingdom, Mary Lyon Centre, Medical Research Council Harwell, Oxfordshire, United Kingdom

Kate Vowell

Affiliation Mary Lyon Centre, Medical Research Council Harwell, Oxfordshire, United Kingdom

Tertius A. Hough

Affiliation Mary Lyon Centre, Medical Research Council Harwell, Oxfordshire, United Kingdom

Lynn Jones

Affiliation Mary Lyon Centre, Medical Research Council Harwell, Oxfordshire, United Kingdom

Paras Pathak

Affiliation Mary Lyon Centre, Medical Research Council Harwell, Oxfordshire, United Kingdom

Hayley E. Tyrer

Affiliation Medical Research Council Mammalian Genetics Unit, Medical Research Council Harwell, Oxfordshire, United Kingdom

Michelle Kelly

Affiliation Medical Research Council Mammalian Genetics Unit, Medical Research Council Harwell, Oxfordshire, United Kingdom

Roger Cox

Affiliation Medical Research Council Mammalian Genetics Unit, Medical Research Council Harwell, Oxfordshire, United Kingdom

Madhuri V. Warren

Affiliation Pathology Diagnostics Ltd, Cambridge, United Kingdom

Jo Peters

Affiliation Medical Research Council Mammalian Genetics Unit, Medical Research Council Harwell, Oxfordshire, United Kingdom

Competing Interests

Co-author MVW is now the director of a commercial company (Pathology Diagnostics Ltd). There are no relevant conflicts relating to employment, consultancy, patents, products in development or marketed products etc. with respect to the work on this animal model, as this is not the subject of the business. MVM is purely an academic collaborator who made her contributions to the phenotying of this mouse model before the advent of her company. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MTC JP. Performed the experiments: MTC KV TAH PP LJ MK JP. Analyzed the data: MTC KV TAH HET MK RC MVW JP. Wrote the paper: MTC MVW JP.